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Staff information

Alex Guenther

Alex Guenther

Atmospheric Measurement & Data Sciences
Laboratory Fellow
Pacific Northwest National Laboratory
PO Box 999
MSIN: K9-24
Richland, WA 99352


Dr. Alex Guenther received a B.S. in Biology with a minor in Computer Science from the University of Puget Sound and an M.S. and PhD in Civil and Environmental Engineering from Washington State University. After finishing a postdoctoral fellowship at the U. Colorado - Boulder in 1990, he was hired as a scientist in the Atmospheric Chemistry Division of NCAR where he was a senior scientist and section head until he joined PNNL in 2013. He was recently awarded the AGU Kaufman award for unselfish cooperation in research and currently serves as the co-chair of the IGBP iLEAPS activity. Dr. Guenther has investigated the role of trace gases and aerosols in biosphere-atmosphere interactions for more than 25 years and is the author of more than 240 peer reviewed journal articles (H index > 60, total citations > 15000). Dr. Guenther is the lead developer of the MEGAN model that is widely used by the scientific and regulatory communities to simulate biogenic emissions for regional air quality and global earth system modeling. He has also conducted numerous laboratory studies and has led more than 40 integrative field studies on six continents in tropical, temperate and boreal ecosystems to provide observations to advance understanding of biogenic emissions and their role in air quality and climate.

PNNL Publications


  • Adam JC, JC Stephens, S Chung, MP Brady, RD Evans, CE Kruger, BK Lamb, ML Liu, CO Stockle, JK Vaughan, K Rajagopalan, J Harrison, CL Tague, A Kalyanaraman, Y Chen, AB Guenther, FY Leung, LYR Leung, AB Perleberg, J Yoder, E Allen, S Anderson, B Chandrasekharan, K Malek, T Mullis, C Miller, T Nergui, J Poinsatte, J Reyes, J Zhu, JS Choate, X Jiang, R Nelson, JH Yoon, GG Yorgey, K Johnson, KJ Chinnayakanhalli, AF Hamlet, B Nijssen, and V Walden. 2015. "BioEarth: Envisioning and developing a new regional earth system model to inform natural and agricultural resource management." Climatic Change 129(3-4):555-571.  doi:10.1007/s10584-014-1115-2
  • Chen Q, DK Farmer, LV Rizzo, T Pauliqueivis, M Kuwata, TG Karl, AB Guenther, JD Allan, H Coe, MO Andreae, U Poeschl, JL Jiminez, P Artaxo, and ST Martin. 2015. "Submicron particle mass concentrations and sources in the Amazonian wet season (AMAZE-08)." Atmospheric Chemistry and Physics 15(7):3687-3701.  doi:10.5194/acp-15-3687-2015
  • Jardine K, AM Yanez-Serrano, J Williams, N Kunert, A Jardine, T Taylor, L Abrell, P Artaxo, AB Guenther, CN Hewitt, E House, AP Florentino, A Manzi, N Higuchi, J Kesselmeier, T Behrendt, PR Veres, B Derstroff, J Fuentes, ST Martin, and MO Andreae. 2015. "Dimethyl sulfide in the Amazon rain forest." Global Biogeochemical Cycles 29(1):19-32.  doi:10.1002/2014GB004969
  • Kim S, AB Guenther, B Lefer, J Flynn, R Griffin, AP Rutter, L Gong, and BK Cevik. 2015. "Potential role of stabilized Criegee radicals in sulfuric acid production in a high biogenic VOC environment." Environmental Science & Technology 49(6):3383-3391.  doi:10.1021/es505793t
  • Kim S, SY Kim, M Lee, H Shim, G Wolfe, AB Guenther, A He, Y Hong, and J Han. 2015. "Impact of isoprene and HONO chemistry on ozone and OVOC formation in a semirural South Korean forest." Atmospheric Chemistry and Physics 15(8):4357-4371.  doi:10.5194/acpd-14-16691-2014


  • Brauer CS, TA Blake, AB Guenther, SW Sharpe, RL Sams, and TJ Johnson. 2014. "Quantitative infrared absorption cross sections of isoprene for atmospheric measurements." Atmospheric Measurement Techniques 7(11):3839-3847.  doi:10.5194/amt-7-3839-2014
  • Acosta Navarro JC, S Smolander, H Struthers, E Zorita, AM Ekman, JO Kaplan, AB Guenther, A Arneth, and I Riipinen. 2014. "Global Emissions of Terpenoid VOCs from Terrestrial Vegetation in the Last Millennium." Journal of Geophysical Research. D. (Atmospheres) 119(11):6867-6885.  doi:10.1002/2013JD021238
  • Gierens R, L Laakso, D Mogensen, V Vakkari, JP Beukes, P Van Zyl, H Hakola, AB Guenther, JJ Pienaar, and M Boy. 2014. "Modelling New Particle Formation Events in the South African Savannah." South African Journal of Science 110(5/6):Article No. 2013-0108.  doi:10.1590/sajs.2014/20130108
  • Gierens R, L Laakso, D Mogensen, V Vakkari, JP Buekes, P Van Zyl, H Hakola, AB Guenther, JJ Pienaar, and M Boy. 2014. "Modelling new particle formation events in the South African savannah." South African Journal of Science 110(5/6):59-70.  doi:10.1590/sajs.2014/20130108
  • Gonzalez N, AK Borg-Karlson, P Artaxo, AB Guenther, R Krejci, B Noziere, and K Noone. 2014. "Primary and secondary organics in tropical Amazonian rainforest aerosols: Chiral analysis of 2-methyltetrols." Environmental Science Processes & Impacts 16(6):1413-1421.  doi:10.1039/C4EM00102H
  • Greenberg J, J Penuelas, AB Guenther, R Seco, A Turnipseed, X Jiang, I Filella, M Estiraste, J Sardans, R Ogaya, J Llusia, and F Rapparini. 2014. "A tethered-balloon PTRMS sampling approach for surverying of landscape-scale biogenic VOC fluxes." Atmospheric Measurement Techniques 7(7):2263-2271.  doi:10.5194/amt-7-2263-2014
  • Harley P, A Eller, AB Guenther, and RK Monson. 2014. "Observations and models of emissions of volatile terpenoid compounds from needles of ponderosa pine trees growing in situ: Controls by light, temperature and stomatal conductance." Oecologia 176(1):35-55.  doi:10.1007/s00442-014-3008-5
  • Jaars K, JP Beukes, PG van Zyl, AD Venter, M Josipovic, JJ Pienaar, V Vakkari, H Aaltonen, H Laakso, M Kulmala, P Tiitta, AB Guenther, H Hellen, L Laakso, and H Hakola. 2014. "Ambient aromatic hydrocarbon measurements at Welgegund, South Africa." Atmospheric Chemistry and Physics 14(13):7075-7089.  doi:10.5194/acp-14-7075-2014
  • Kim S, T VandenBoer, C Young, T Riedel, JA Thornton, B Swarthout, B Sive, BM Lerner, J Gilman, C Warneke, JM Roberts, AB Guenther, N Wagner, WP Dube, E Williams, and S Brown. 2014. "The Primary and Recycling Sources of OH During the NACHTT-2011 Campaign: HONO as an Important OH Primary Source in the Wintertime." Journal of Geophysical Research. D. (Atmospheres) 119(11):6886-6896.  doi:10.1002/2013JD019784
  • Liu ML, K Rajagopalan, SH Chung, X Jiang, JH Harrison, T Nergui, AB Guenther, C Miller, J Reyes, CL Tague, JS Choate, E Salathe, CO Stockle, and JC Adam. 2014. "What is the importance of climate model bias when projecting the impacts of climate change on land surface processes?" Biogeosciences 11(10):2601-2622.  doi:10.5194/bg-11-2601-2014
  • Marais EA, D Jacob, AB Guenther, K Chance, TP Kurosu, JG Murphy, CE Reeves, and H Pye. 2014. "Improved Model of Isoprene Emissions in Africa using Ozone Monitoring Instrument (OMI) Satellite Observations of Formaldehyde: Implications for Oxidants and Particulate Matter." Atmospheric Chemistry and Physics 14(15):7693-7703.  doi:10.5194/acp-14-7693-2014
  • Misztal P, TG Karl, R Weber, HH Jonsson, AB Guenther, and AH Goldstein. 2014. "Airborne flux measurements of Biogenic Isoprene over California." Atmospheric Chemistry and Physics 14(19):10631-10647.  doi:10.5194/acp-14-10631-2014
  • Nakashima Y, S Kato, J Greenberg, P Harley, TG Karl, A Turnipseed, E Apel, AB Guenther, J Smith, and Y Kajii. 2014. "Total OH reactivity measurements in ambient air in a southern Rocky Mountain ponderosa pine forest during BEACHON-SRM08 summer campaign." Atmospheric Environment 85:1-8.  doi:10.1016/j.atmosenv.2013.11.042
  • Ortega J, A Turnipseed, AB Guenther, TG Karl, DA Day, D Gochis, JA Huffman, AJ Prenni, EJ Levin, SM Kreidenweis, PJ DeMott, Y Tobo, EG Patton, A Hodzic, YY Cui, P Harley, RS Hornbrook, EC Apel, RK Monson, AS Eller, JP Greenberg, M Barth, P Campuzano-Jost, BB Palm, JL Jiminez, AC Aiken, MK Dubey, C Geron, J Offenberg, MG Ryan, PJ Fornwalt, SC Pryor, FN Keutsch, JP DiGangi, AW Chan, AH Goldstein, GM Wolfe, S Kim, L Kaser, R Schnitzhofer, A Hansel, C Cantrell, RL Mauldin, and JN Smith. 2014. "Overview of the Manitou Experimental Forest Observatory: site description and selected science results from 2008 to 2013." Atmospheric Chemistry and Physics 14(12):6345-6367.  doi:10.5194/acp-14-6345-2014
  • Park R, SK Hong, HA Kwon, S Kim, AB Guenther, JH Woo, and CP Loughner. 2014. "An evaluation of ozone dry deposition simulations in East Asia." Atmospheric Chemistry and Physics 14(15):7929-7940.  doi:10.5194/acp-14-7929-2014
  • Potosnak M, L LeStourgeon, SG Pallardy, KP Hosman, L Gu, T Karl, C Geron, and AB Guenther. 2014. "Observed and modeled ecosystem isoprene fluxes from an oak-dominated temperate forest and the influence of drought stress." Atmospheric Environment 84:314-322.  doi:10.1016/j.atmosenv.2013.11.055
  • Sindelarova K, C Granier, I Bouarar, AB Guenther, S Tilmes, T Stavrakou, JF Muller, U Kuhn, P Stefani, and W Knorr. 2014. "Global data set of biogenic VOC emissions calculated by the MEGAN model over the last 30 years." Atmospheric Chemistry and Physics 14(17):9317-9341.  doi:10.5194/acp-14-9317-2014
  • Situ S, X Wang, AB Guenther, Y Zhang, X Wang, M Huang, Q Fan, and Z Xiong. 2014. "Uncertainties of isoprene emissions in the MEGAN model estimated for a coniferous and broad-leaved mixed forest in Southern China." Atmospheric Environment 98:105-110.  doi:10.1016/j.atmosenv.2014.08.023
  • Smolander S, Q He, D Mogensen, L Zhou, J Back, T Ruuskanen, S Noe, AB Guenther, H Aaltonen, M Kulmala, and M Boy. 2014. "Comparing three vegetation monoterpene emission models to measured gas concentrations with a model of meteorology, air chemistry and chemical transport." Biogeosciences 11(19):5425-5443.  doi:10.5194/bg-11-5425-2014
  • Stavrakou T, JF Muller, M Bauwens, I De Smedt, VM Roozendael, AB Guenther, M Wild, and X Xia. 2014. "Isoprene emissions over Asia 1979-2012 : impact of climate and land use changes." Atmospheric Chemistry and Physics 14(9):4587-4605.  doi:10.5194/acp-14-4587-2014
  • Wolfe GM, C Cantrell, S Kim, RL Mauldin, TG Karl, P Harley, A Turnipseed, W Zheng, FM Flocke, EC Apel, RS Hornbrook, SR Hall, K Ullmann, SB Henry, JP DiGangi, ES Boyle, L Kaser, R Schnitzhofer, A Hansel, M Graus, Y Nakashima, Y Kajii, AB Guenther, and FN Keutsch. 2014. "Missing Peroxy Radical Sources within a Summertime Ponderosa Pine Forest." Atmospheric Chemistry and Physics 14(9):4715-4732.  doi:10.5194/acp-14-4715-2014
  • You Y, VP Kanawade, JA de Gouw, AB Guenther, S Madronich, MR Sierra-Hernandez, M Lawler, JN Smith, S Takahama, G Ruggeri, A Koss, K Olson, K Baumann, RJ Weber, A Nenes, H Guo, ES Edgerton, L Porcelli, WH Brune, AH Goldstein, and SH Lee. 2014. "Atmospheric Amines and Ammonia Measured with a Chemical Ionization Mass Spectrometer (CIMS)." Atmospheric Chemistry and Physics 14(22):12181-12194.  doi:10.5194/acp-14-12181-2014
  • Yu H, J Ortega, JN Smith, AB Guenther, VP Kanawade, Y You, Y Liu, KP Hosman, T Karl, R Seco, C Geron, SG Pallardy, L Gu, J Mikkila, and SH Lee. 2014. "New Particle Formation and Growth in an Isoprene-Dominated Ozark Forest: From Sub-5 nm to CCN-Active Sizes." Aerosol Science and Technology 48(12):1285-1298.  doi:10.1080/02786826.2014.984801
  • Zhang H, Z Zhang, T Cui, YH Lin, NA Bhathela, J Ortega, D Worton, AH Goldstein, AB Guenther, JL Jimenez, A Gold, and JD Surratt. 2014. "Secondary Organic Aerosol Formation from 2-Methyl-3-Buten-2-ol Photooxidation: Evidence of Acid-Catalyzed Reactive Uptake of Epoxides." Environmental Science & Technology Letters 1(4):242-247.  doi:10.1021/ez500055f


  • Fares S, R Schnitzhofer, J Xiaoyan, AB Guenther, A Hansel, and F Loreto. 2013. "Observations of Diurnal to Weekly Variations of Monoterpene-Dominated Fluxes of Volatile Organic Compounds from Mediterranean Forests: Implications for Regional Modeling." Environmental Science & Technology 47(19):11073-11082.  doi:10.1021/es4022156
  • Frost GJ, P Middleton, L Tarrason, C Granier, AB Guenther, B Cardenas, H Denier van der Gon, G Janssens-Maenhout, JW Kaiser, T Keating, Z Klimont, JF Lamarque, C Liousse, S Nickovic, T Ohara, M Schultz, U Skiba, and Y Wang. 2013. "New Directions: GEIA's 2020 Vision for Better Air Emissions Information." Atmospheric Environment 81(78):710-712.  doi:10.1016/j.atmosenv.2013.08.063
  • Kaser L, T Karl, AB Guenther, M Graus, R Schnitzhofer, A Turnipseed, L Fischer, P Harley, M Madronich, D Gochis, FN Keutsch, and A Hansel. 2013. "Undisturbed and disturbed above canopy ponderosa pine emissions: PTR-TOF-MS measurements and MEGAN 2.1 model results." Atmospheric Chemistry and Physics 13(23):11935-11947.  doi:10.5194/acp-13-11935-2013
  • Mak JE, L Su, AB Guenther, and TG Karl. 2013. "A novel Whole Air Sample Profiler (WASP) for the quantification of volatile organic compounds in the boundary layer." Atmospheric Measurement Techniques 6(10):2703-2712.  doi:10.5194/amt-6-2703-2013
  • Situ S, AB Guenther, XJ Wang, X Jiang, A Turnipseed, Z Wu, J Bai, and X Wang. 2013. "Impacts of seasonal and regional variability in biogenic VOC emissions on surface ozone in the Pearl River Delta region, China." Atmospheric Chemistry and Physics 13(23):11803-11817.  doi:10.5194/acp-13-11803-2013
  • Unger N, K Harper, Y Zheng, NY Kiang, I Aleinov, A Arneth, G Schurgers, C Amelynck, AH Goldstein, AB Guenther, B Heinesch, CN Hewitt, T Karl, Q Laffineur, B Langford, KA McKinney, P Misztal, M Potosnak, J Rinne, S Pressley, N Schoon, and D Serca. 2013. "Photosynthesis-dependent isoprene emission from leaf to planet in a global carbon-chemistry-climate model." Atmospheric Chemistry and Physics 13(20):10243-10269.  doi:10.5194/acp-13-10243-2013

Selected Publications


  • Guenther AB, X Jiang, CL Heald, T Sakulyanontvittaya, T Duhl, L K. Emmons and X Wang. 2012. "The Model of Emissions of Gases and Aerosols from Nature version 2.1 (MEGAN2.1): an extended and updated framework for modeling biogenic emissions." Geoscientific Model Development 5(6):1471-1492.


  • Guenther A, M Kulmala, A Turnipseed, J Rinne and T Sunni. 2011. "Integrated land ecosystem - atmosphere processes study (iLEAPS) assessment of global observational networks." Boreal Environment Research 16:321-336.

  • Hewitt, CN, K Ashworth, A Boynard, A Guenther, B Langford, AR MacKenzie, PK Misztal, E Nemitz, SM Owen, M Possell, TAM Pugh, AC Ryan and O Wild. 2011. "Ground-level ozone influenced by circadian control of isoprene emissions." Nature Geosciences 4(10):671-674.


  • Karl, T, P Harley, L Emmons, B Thornton, A Guenther, C Basu, A Turnipseed and K Jardine. 2010.  "Efficient Atmospheric Cleansing of Oxidized Organic Trace Gases by Vegetation." DOC:10.1126/science.1192534, Science 3(30): 816-819.


  • Guenther, A, T Karl, P Harley, C Wiedinmyer, P Palmer and C Geron. 2006. "Estimates of global terrestrial isoprene emissions using MEGAN (Model of Emissions of Gases and Aerosols from Nature)." Atmospheric Chemistry and Physics 6:3181-3210. DOI: 10.5194/acp-6-3181-2006.


  • Guenther, A, CN Hewitt, D Erickson, R Fall, C Geron, T Graedel, P Harley, L Klinger, M Lerdau, W McKay, T Pierce, B Scholes, R Steinbrecher, R Tallamraju, J Taylor and P Zimmerman. 1995. "A global model of natural volatile organic compound emissions." Journal of Geophysical Research 100: 8873-8892. DOI: 10.1029/94JD02950.


  • Guenther, A, P Zimmerman, P Harley, R Monson and R Fall. 1993. "Isoprene and monoterpene emission rate variability: Model evaluation and sensitivity analysis." Journal of Geophysical Research 98:12609-12617. DOI: 10.1029/93JD00527.

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